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In page Coprolite:

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The fossil hunter Mary Anning noticed as early as 1824 that "bezoar stones" were often found in the abdominal region of ichthyosaur skeletons found in the Lias formation at Lyme Regis.[1] She also noted that if such stones were broken open they often contained fossilized fish bones and scales as well as sometimes bones from smaller ichthyosaurs. It was these observations by Anning that led the geologist William Buckland to propose in 1829 that the stones were fossilized feces and name them coprolites. Buckland also suspected that the spiral markings on the fossils indicated that ichthyosaurs had spiral ridges in their intestines similar to those of modern sharks, and that some of these coprolites were black with ink from swallowed belemnites.[2]